Harwood F. Byrnes/Richard B. Sewall Teaching Prize, given by Yale College for a long period of service that inspired a great number of students and consistently fostered learning both inside and outside the classroom, 2021.
Honorary Doctorate in Natural Sciences, University of Zurich, 2015
DeVane Medal for distinction in undergraduate teaching, 2011. Laudatio 1, Laudatio 2
Distinguished Fellow, European Society for Evolutionary Biology, 2007
Honorary Membership in the Swiss Zoological Society, 2005. Laudatio
Fellow, AAAS 1987